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College and Graduate School of Arts and Sciences

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Catalog of Courses for Biology

BIOL 1040
The DNA Revolution

Imagine a world where your DNA is sequenced for free and any human gene can be altered at will. The goal of this course is to address the question: can our society be better prepared for this transformation in science? Is genetic privacy achievable or genetic discrimination avoidable? Who owns your genes? Do your genes drive your medical future? Classes involve student perspectives and discussions with experts in science, policy, ethics and law.

BIOL 1050
Genes and Citizens

Genetics and Genomics form the basis for much of modern biology and the future of medical practice. A basic understanding of them is important for people to be able to evaluate the science behind many issues both public and private. Genetics and Genomics and some of the ways they confront and inform modern life will be covered in a way that is accessible to non-scientists.

BIOL 1060
Principles of Nutrition

Paleo or South Beach? Are supplements wise? Together we will investigate advertising claims, discover & evaluate nutritional resources, discuss public policies & food industry regulations, search through data from epidemiological studies and read clinical cases. To do this, we will delve deep into the physiological workings of the gastrointestinal tract, as well as the molecular metabolic pathways that cells and tissues need to survive & thrive.

Course was offered:  Spring 2017 · Spring 2016 · Spring 2015
BIOL 1080
Behavioral Neurobiology

Ecolocation in bats, development of learning in songbirds, paralytic goats and toxic fish. In this course, we'll examine these and other examples from nature to model the fundamental properties of neurons and the neural circuits that underlie various aspects of animal behavior. Building an understanding of the structure & function of the nervous system will include consideration of the evolutionary and developmental emergence of its properties.

Course was offered:  Spring 2017 · Spring 2016
BIOL 1210
Human Biology and Disease

Introduces how the human body works using basic biological principles. Uses disease as a lens to develop healthcare literacy and to understand fundamental healthcare decisions. This course provides tools to help make informed choices as voters and consumers.

BIOL 150
Special Topics in Biology

Special Topics in Biology.

Course was offered:  Summer 2020
BIOL 1559
New Course: BIOL

New course in the subject of Biology.

BIOL 2010
Intro Bio:Cell Biol & Genetics

BIOL 2100 is one of two semester courses that together provide an intensive introduction to biology for prospective Biology majors and pre-health (med, vet, dental) students. This course focuses on the fundamentals of cell biology and genetics with an emphasis on classical and modern experimental approaches. Lecture topics and concepts are reinforced and extended during once-weekly laboratory/small group discussions.

BIOL 2020
Intro Biol:Orgnsm & Evol Biol

BIOL 2200 is one of two semester courses that together provide an intensive introduction to biology for prospective Biology majors and pre-health (med, vet, dental) students. This course focuses on evolution, physiology and development. Lecture topics and concepts are reinforced and extended during once-weekly laboratory/small group discussions. The Introductory courses are not sequenced and may be taken in either order.

BIOL 2030
Introductory Biology Lab

An investigative experience illustrating modern methods of studying genes and proteins including techniques of DNA isolation, separation, cloning, sequencing, creating recombinant DNA, and using bioinformatics tools. Prerequisite: Limited to 2nd, 3rd, 4th year students who have completed BIOL2010

BIOL 2040
Intro Biol Lab II

Studies life forms, from simple to complex organization, demonstrating the unique properties of living organisms. Exercises focus on evolution, physiology and development. Prerequisite: Limited to 2nd, 3rd, 4th year students who have completed BIOL2020

BIOL 2100
IntroBio w/Lab:Cell & Genetics Offered Spring 2026

BIOL 2100 is one of two semester courses that together provide an intensive introduction to biology for prospective Biology majors and pre-health (med, vet, dental) students. This course focuses on the fundamentals of cell biology and genetics with an emphasis on classical and modern experimental approaches. Lecture topics and concepts are reinforced and extended during once-weekly laboratory/small group discussions.

BIOL 2200
Intro Biol:Orgnsm & Evol Biol Offered Spring 2026

BIOL 2200 is one of two semester courses that together provide an intensive introduction to biology for prospective Biology majors and pre-health (med, vet, dental) students. This course focuses on evolution, physiology and development. Lecture topics and concepts are reinforced and extended during once-weekly laboratory/small group discussions. The Introductory courses are not sequenced and may be taken in either order.

BIOL 2757
Science Writing

Writing is fundamental to the practice of science. We write about individual organisms, ecosystems, and patterns, to record our findings and to reach broader audiences. This course explores diverse writing styles to improve student communication both inside scientific communities and to the public. Students will be inspired by their experiences at MLBS and by prominent nature and science writers to create a variety of written works.

BIOL 2900
UTA Teaching Methods

This STEM teaching course will help Undergraduate TAs integrate learning theory and effective student engagement practices into their teaching. UTAs will participate in guided discussions to relate recommendations from the education literature to their classroom experiences. Assignments will include learning activities, such as teaching observations & reflections, and designing interventions to assist students with difficult topics/skills.

Course was offered:  Fall 2018 · Fall 2017 · Fall 2016
BIOL 3000

Examines the fundamental principles of eukaryotic cell biology at the molecular level. Topics include: structure and function of the plasma membrane, transport of small molecules, ions and macromolecular complexes across membranes, protein trafficking, the cytoskeleton, signal transduction pathways, and the control of cell division and cellular proliferation. Prerequisites: completion of BIOL 2100 or BME 2104 and either CHEM 1410,1420, or CHEM 1810,1820. NOT repeatable if passing grade received.

BIOL 3010
Genetics and Molecular Biology Offered Spring 2026

What makes humans different from fruit flies? Why does your brain have neurons and not liver cells? This course is all about the answer to these questions: It's the genes! This course covers the chemical make-up of genes, how they're passed on through generations, how they're expressed and how that expression is regulated, how disruption in the structure and expression of genes arise and how those disruptions lead to cellular defects and disease. Prerequisite: Must have completed BIOL 2100 or BME 2104 and either CHEM 1410 or CHEM 1810.  BIOL 3010 is not repeatable. 

BIOL 3020
Evolution and Ecology

Examines the mechanisms of evolutionary change, with an emphasis on the genetic and evolutionary principles needed to understand the diversification of life on earth.  Covers the ecology of individuals and population dynamics.  Major topics include the genetics and ecology of natural populations, adaptation, molecular evolution and macroevolution, and the application of evolutionary and ecological concepts to conservation biology.  Required for all Biology majors. Prerequisite: Must have completed BIOL 2200. BIOL 3020 is not repeatable.

BIOL 3030

Biochemistry underlies nearly every biological process, from environmental science to medicine. When living systems are in chemical and energetic balance, organisms thrive. When they're out of balance, as in disease or unpredictable environments, life is compromised. This course will explain how simple chemical and physical principles apply to the major classes of biological macromolecules that maintain life. Prerequisite: BIOL 2010 or BIOL 2100 or BME 2104 and BIOL 2020 or BIOL 2040 and either CHEM 2410 or CHEM 1820

BIOL 3040
Develop and Regen Biol Offered Spring 2026

Are developmental biology and regenerative biology one and the same? Throughout this course, we will emphasize both classical and modern experimental approaches that have been used to unravel the genetic, molecular and celluar mechanisms of development. Additionally, the practical value of understanding development is enormous, and the relationship between embryology and clinical applications will be a theme that runs throughout the course.

BIOL 3050
Introduction to Neurobiology

Analyzes the concepts of general neurobiology, including basic electrophysiology and electrochemistry, origin of bioelectric potentials, sensory, motor, integrative and developmental neurobiology, and conceptual models of simple learning. Prerequisites: BIOL 2100 (or BME 2104) and BIOL 2200. Recommended:  Prior completion of BIOL 3000

BIOL 3090
Infectious Disease

Infectious disease impacts every human, plant and animal on earth. What is the most deadly disease in human history? What is killing our ocean's turtles? Why is Zika so scary? We will explore questions related to the biology, transmission, and pathogenicity of infectious agents across the world. We will also place special emphasis on what it takes to successfully control an infectious disease.

BIOL 3120
Microbiology Genomics Era

Microbes rule. In this course, we will explore how microbes rule the world and how genomics has revolutionized the way we study them. Fundamental principles of microbiology, together with the basics of genomics will be introduced. Topics include microbial cell structure, metabolism, genetics, microbial diversity and ecology, epidemiology, genome sequencing technologies and bioinformatics. Prerequisites: Must have completed BIOL 2010 or BIOL 2100 or BME 2104 and BIOL 2020 or BIOL 2200

BIOL 3140
Biology of Aging

This interdisciplinary course will explore our current knowledge of the biology of aging in populations of plants and animals, including humans. Topics include demographic trends across species; analysis of why organisms age in the context of evolutionary theories; analysis of how organisms age in the context of cellular and physiological theories; and the genetic basis of longevity. Prerequisites: BIOL 2010 and 2020.

Course was offered:  Fall 2019 · Fall 2014
BIOL 3150
Microbiology Laboratory

An introduction to microorganisms and to basic microbiological principles through laboratory experimentation. Emphasis is on the structure, physiology and genetics of bacteria and bacterial viruses. Prerequisite: BIOL 3120

BIOL 3170
Introduction to Neurobiology

Analyzes the concepts of general neurobiology, including basic electrophysiology and electrochemistry, origin of bioelectric potentials, sensory, motor, integrative and developmental neurobiology, and conceptual models of simple learning. Prerequisites: BIOL 2100 (or BME 2104) and BIOL 2200. Recommended:  Prior completion of BIOL 3000

Course was offered:  Fall 2014 · Fall 2013
BIOL 3180
Plants and Society Offered Spring 2026

This is an introductory course that takes a multidisciplinary approach to studying the relationship between plants and people. The course focuses on providing students foundational information on the growth, development, physiology and genetics of plants and explores the connection between plants and people by looking at the use of plants as sources of food, shelter, medicinals and manufactured goods.

BIOL 3230
Animal Physiology

Focuses on selected vertebrate organ systems; considers other systems where relevant. Prerequisite: BIOL 2010 and 2020.

BIOL 3240
Introduction to Immunology Offered Spring 2026

Studies the genetics and cell biology of the vertebrate immune system, with a focus on adaptive immunity. Classic and current experimental systems are emphasized. Prerequisite: Must have completed or be currently taking BIOL 2010 or BIOL 2100 or BME 2104

BIOL 3250
Intro to Animal Behavior

An introduction to comparative studies of animal behavior from neuroethological and evolutionary prospectives. The first deals with proximate causes of behavior, with emphasis on motor, sensory and central aspects of the nervous system. The second deals with ultimate causes, with emphases on natural selection, natural history, and adaptive aspects of behavior.

BIOL 3260
Editing Genes and Genomes Offered Spring 2026

Genome databases contain a wealth of information that enable us to answer myriad questions in biology. Working with genome data requires foundational knowledge in molecular genetic concepts, as well as technical knowledge of how to read and analyze sequence data. This class will provide students with the skills to understand genomic data and its applications in biology and medicine.

BIOL 3270
Microbiology with Lab Offered Spring 2026

Microbes rule. In this course, we will explore how microbes rule the world and how genomics has revolutionized the way we study them. Fundamental principles of microbiology will be introduced. Topics include microbial cell structure, metabolism, genetics, diversity, evolution and infectious disease. Laboratory work will complement lecture topics and cover the core themes & concepts, as recommended by the American Society of Microbiology.

BIOL 3360
Biological Therapy of Cancer

This seminar course revolves around student-led presentations of primary literature in the field of cancer therapy using novel approaches including immunotherapies. Objectives include providing the student with significant exposure to primary literature and the development of critical thinking skills. Prerequisite: BIOL 3240.

Course was offered:  Spring 2015 · Spring 2014
BIOL 3410
Human Anatomy and Physiology I

This course, the first in a two-course sequence, is an introduction to the structure and function of the human body. Review of the structure and physiology of cells and tissues leads to in-depth study of the musculoskeletal and nervous systems. Control mechanisms and the contributions of each system to overall homeostasis are emphasized.

BIOL 3420
Human Anatomy & Physiology II Offered Spring 2026

This course, the second in a two-course sequence, examines structures and functions of the endocrine, cardiovascular, urogenital, respiratory, renal, gastrointestinal and reproductive systems. Control mechanisms and functional integration of these systems in overall homeostasis is emphasized.

BIOL 3430
Physio Mech in Health Disease Offered Spring 2026

Emphasis on the functions and integration of human nervous, cardiovascular, respiratory, digestive, and renal systems in maintaining homeostasis, and by extension, health. Normal function, from cells to organs, of each system provides a foundation for study of mechanisms that lead to dysfunction and the identification of potential therapeutic targets and strategies.

Course was offered:  Spring 2026 · Spring 2025
BIOL 3450
Biodiversity and Conservation

Introduction to the fundamental principles of conservation biology (e.g., global species numbers, value of biodiversity, causes of extinction, genetic diversity, island biogeography, priority setting) and current topics of debate (including zoo versus field conservation, effects of global change on species extinction). Conservation case studies will allow students to judge the relevance of biological theory to practical problems in conservation.

BIOL 3559
New Course: BIOL
BIOL 3585
Selected Topics in Biology

Tutorial or seminar course that allows intensive study of the literature in a particular area of biology under the guidance of a Biology faculty member.

Course was offered:  Fall 2014 · Fall 2013
BIOL 3710
The Biology of Stress

What exactly is stress? When is it a good thing; when & why does it become damaging? In this course, we will study how the body responds to physical and psychological stressors. And, we will examine how the physiological mechanisms by which the hypothalamic pituitary adrenal axis and corticosteroids mediate both positive and negative effects of stress. Understanding of these mechanisms, we can consider how best to prevent damage from stress.

Course was offered:  Fall 2018 · Fall 2017
BIOL 3900
Independent Readings

Tutorial or seminar course that allows intensive study of the literature in a particular area of biology under the guidance of a Biology faculty member.

BIOL 4005
Functional Genomics Laboratory

This course introduces students to scientific-based discovery of how molecular dysfunction leads to disease. It also exposes them to the most current tools used in biomedical research to find novel genes and compounds that could help treat human disease. The course includes discovery-oriented lab, workshops, and lectures. Prerequisite: BIOL3000 and BIOL3010

BIOL 4011
Homeostasis:Wisdom of the Body

The human body maintains stable energy levels, hydration, and temperature despite the challenges of ever-changing external environment, a process known as homeostasis. This course explores biological models and mechanisms of homeostasis, including how survival needs are monitored and met through changes in behavior and physiology. Students will gain a state-of-the-art perspective on homeostatic biology and its research methods and technology.

Course was offered:  Spring 2024 · Spring 2022 · Spring 2021
BIOL 4012
Disease Ecology and Evolution Offered Spring 2026

In this course, we'll dive into our current understanding of the evolution and ecology of parasitic interactions through primary literature, modeling, and experimental design. Throughout, we will focus on generating and testing hypotheses, evaluating theoretical models with evidence, drawing parallels between diverse domains of life, and connecting evolutionary and ecological ideas to today's past, present, and future epidemics.

Course was offered:  Spring 2026 · Fall 2024 · Fall 2021
BIOL 4013
Stem Cells

The course will deep dive into what stem cells are, what they do, where and how they function, and how we can use stem cells in the clinic to repair damaged tissue and restore tissue function. The course will consist of a series of lectures and student run discussions related to current scientific literature.

Course was offered:  Spring 2022
BIOL 4014
Cell Origin-Animal Diversity Offered Spring 2026

Animals are incredibly diverse, but they all evolved from the same single-celled ancestor that lived hundreds of millions of years ago. This course takes a cell-biological approach to explore key questions in animal evolution such as the origins of multicellularity and differentiation. Students will gain a cutting-edge perspective on current research that integrates cell, developmental, and evolutionary biology to explore animal origins.

BIOL 4015
Neur Dev Lab

Neural stem cells proliferate throughout development to generate the immense diversity of neuronal cell types present in our adult brains. What are the signals that drive neural stem cells to proliferate & what are the signals that terminate stem cell divisions once development is complete? Using Drosophila we will investigate these questions and address specifically the role of nutrition in regulating profileration of the stem cell population. Prerequisite: BIOL 3000, BIOL 3010

BIOL 4016
Precision Medicine

This course addresses the impact of the human genome project on understanding human genetic disease, focusing on the invaluable role for animal models of diseases in augmenting evaluation of genomic information to develop strategies for precision medicine. Animal models are an invaluable asset in reaching this goal because they allow experimental manipulations that go far beyond what is possible in human patients.

Course was offered:  Spring 2025 · Spring 2023 · Spring 2022
BIOL 4017
The Immortal Germ Line Offered Spring 2026

One of the most important characteristics of life is the ability to reproduce. In order to produce new life, multicellular organisms evolved specialized cells whose only purpose is reproduction ¿ the germ cells. Germ cells are the only cells that persist from one generation to the next and are often called immortal. We will decipher how these totipotent stem cells function in order to faithfully create the next generation of organisms.

Course was offered:  Spring 2026 · Spring 2025
BIOL 4018
Microbial Genome Sequencing

Microbes rule. This course will teach microbial genomics using the cutting edge next-generation DNA sequencing technology and its applications to study microbes around us. Topics covered include microbial genomics, DNA sequencing and sequence analysis.

Course was offered:  Fall 2025 · Fall 2023 · Fall 2022
BIOL 4019
Psychopharmacology of Plants

This course begins with discussion of pharmacological principles and normal function of the nervous and endocrine system. As we continue, we will describe how exogenous substances derived from plants (like drugs) impact the nervous system to restore normal or near-normal function, or alter normal function, in humans. The use of agents from plants in the alleviation of depression and anxiety will be emphasized.

Course was offered:  Fall 2025 · Fall 2024 · Fall 2023
BIOL 4020
Computational Evolutionary Bio

The evolutionary history of a population can be studied by examining patterns of genetic variation among individuals. Using information about genetic variation, we can infer historical evolutionary events like migration and adaptation. In this lab course, you will learn to utilize genomic data to conduct evolutionary inference. We will learn fundamentals of population genetics, bioinformatic skills, and research methods applied to real short-read sequencing data.

Course was offered:  Fall 2025 · Fall 2024
BIOL 4021
Adv in Regenerative Biology

This course will provide an in-depth exploration of the field of regenerative biology, focusing on the molecular and cellular mechanisms underlying tissue regeneration and repair in animals. We will explore the cellular basis of different types of regeneration and search for shared molecular mechanisms. With an eye towards the future, we¿ll also explore the implications for advancing regenerative medicine.

Course was offered:  Fall 2025
BIOL 4030
The Study of Neuroscience Lab Offered Spring 2026

This course introduces students to foundational and modern neuroscience research techniques through laboratory investigations. Neuroscience is explored at the molecular, systems, and behavioral level. Topics covered include neuroanatomy, electrophysiology, biopotentials, development, histology, and microscopy. An emphasis is placed on structure-function relationships, experimental design, and application of techniques in research.

Course was offered:  Spring 2026 · Fall 2025
BIOL 4040
Laboratory in Cell Biology

Introduces students to experimental approaches, including mammalian cell culture, gel electrophoresis, western blotting and immunofluorescence microscopy, that are used to study both normal and pathological processes at the level of individual cells. The biological theme of the course will be Alzheimer's disease (AD) and related neurodegenerative disorders. One laboratory lecture and one afternoon laboratory per week. Prerequisite: BIOL 3000

BIOL 4045
Neurodegenerative Diseases

This course for advanced undergrads will focus mainly on research about Alzheimer's disease, and will meet once/week for 3 hours. The first 3 weeks will be primarily didactic, and the remainder of the course will be a "journal club" in which primary research paper discussions will be led by teams of students. Assessments will be based on how well students lead and participate in discussions, and on exams.

BIOL 4050
Developmental Biology

This course will cover advanced principles of developmental biology and how embryonic developmental pathways impinge on human disease. Topics will include congenital organ related disease, stem cell biology and its therapeutic applications, regenerative medicine and the impact of environmental factors on disease.

BIOL 4060
Organ Dev Tissue Eng

Why do most of our adult body tissues have limited regenerative capacity? How can terminally diseased organs be replaced? This course will cover the cellular mechanisms that regulate animal tissue formation, regeneration and repair in vivo. Students will gain insights into the opportunities, limitations, and risks of tissue engineering in vitro, as an emerging research field that may lead to revolutionary organ replacement strategies. Prerequisite: BIOL 3000

Course was offered:  Spring 2015
BIOL 4070
Developmental Biology Lab Offered Spring 2026

The goal of this course is to provide an original, unknown outcome research experience in developmental biology. After training in basic methods and descriptions of selected research problems, students form teams and investigate a problem of their choosing. Team members work together in the lab, but each writes an independent research proposal, a notebook, and a final project report on which they are graded. Recommended prerequisites: BIOL 3000 and 3010.

BIOL 4080
Advanced Hormones and Behavior

From plants to humans, hormones shape various aspects of organismal form and behavior over contemporary and evolutionary time. Delve into endocrine pathways, hormones' influence on development, and their role in coordinating responses to environmental and physiological stimuli. Gain a deep understanding endocrinology theories, concepts, and methods, and the ability to critically evaluate hormonal impacts on ecosystem and human health.

Course was offered:  Fall 2024
BIOL 4090
Environmental Public Health

This is an interdisciplinary exploration of environmental public health issues. Students develop and research topics, lead small group discussions, give oral presentations, and write papers. Scope of student research in topic development includes env. science, ecology, epidemiology, toxicology, pathophysiology, gene-environment interactions, directions in clinical and translational research, and environmental and biomedical policy development.

BIOL 4100
Management of Forest Ecosystem

An ecosystem course that treats the ecology of forests and consequences of forest processes in natural and managed systems. The class emphasizes the "pattern & process" concept that is the central theme in modern vegetation sciences at increasing scales: from form/function of leaves and other parts of trees through population, community and landscape ecology to the role of forests in the global climate and carbon-cycling. It is recommended to take EVSC 3200 prior to enrollment in this class.

BIOL 4120
When Good Cells Go Bad

This course will cover topics related to major neurodegenerative diseases including Multiple Sclerosis (MS), Muscular Dystrophy (MD), Peripheral Nerve Sheath Tumor (Neurofibrosarcoma) and Malignant peripheral nerve sheath tumor (MPNST). Topics related to pathology and molecular mechanism of diseases, possible drug discovery targets, and therapeutic discovery approaches will be emphasized. Prerequisites: BIOL 3000 and BIOL 3010.

Course was offered:  Fall 2014 · Fall 2013
BIOL 4130
Population Ecol & Consv Biol

The mathematical foundations of population dynamics and species interactions as applied to population and community ecology and problems in conservation biology. One semester of calculus is recommended. Prerequisite: BIOL 3020 or EVSC 3200

BIOL 4135
Biology of Aging

Aging is an evolutionary paradox because it decreases physiological function and increases the risk of mortality, yet aging persists in most species. We will explore the theories of aging and the diversity of the patterns of aging across species from flies to plants to humans. We will use the primary literature in the fields of evolution, genetics and cell biology to gain a comprehensive understand of the latest advances in this field.

Course was offered:  Fall 2021 · Fall 2020 · Fall 2016
BIOL 4140
NextGen Sequencing and Apps

Students will learn the next generation sequencing technologies and explore their applications in the studies of evolution and ecology. This course is a lecture and journal club format where primary scientific literature will be discussed. Students will also learn basic bioinformatic skills. Prerequisite: BIOL 3020

Course was offered:  Fall 2013
BIOL 4150
Evolution of Sex

Despite the many benefits of asexual reproduction, the vast majority of eukaryotic organisms reproduce sexually. How sex evolved, and how it persists despite its many associated costs, are major unanswered questions in biology. We will explore the diversity of sexual reproduction and associated evolutionary phenomena with a focus on critically evaluating current research and theory in this field. Prerequisite: BIOL 3020 or permission from Instructor

BIOL 4180
Behavioral Ecology

Behavioral ecology explores the evolutionary analysis and explanations for the diversity of animal behavior, including foraging decisions, altruism, cooperation, mate choice, group living, parental care and range of other sociobiological phenomena. Prerequisite: BIOL 3020.

Course was offered:  Fall 2014 · Fall 2013
BIOL 4190
Biological Clocks

Introduces biological timekeeping as used by organisms for controlling diverse processes, including sleep-wakefulness cycles, photoperiodic induction and regression, locomotor rhythmicity, eclosion rhythmicity, and the use of the biological clock in orientation and navigation. Prerequisite: BIOL 3000 or 3010 or 3020

BIOL 4210
Genome Sciences

This course will chronicle the meteoric rise in our ability to collect DNA sequence data & reconstruct genomes, and how this contributes to understanding evolution & the genetic basis of traits, including disease. Discussions with leading experts in science, policy or law will allow students to consider the promises & limitations of genomic research, as well as the future societal impact of having nearly ubiquitous genetic information. Prerequisite: BIOL 3010 and BIOL 3020

Course was offered:  Spring 2015
BIOL 4220
Systems Biology

An introduction to a new research paradigm that focuses on the systematic study of complex interactions at the molecular, network and genomic level. This course will review state-of-the-art high throughput techniques and modeling methods used to obtain, integrate and analyze complex data from biological systems. This course will be a combination of text based lectures and discussions of the current literature pertinent to Systems Biology. Prerequisites: BIOL 3010. Also recommended is BIOL 3000

Course was offered:  Spring 2014
BIOL 4230
Bioinformatics

The Genome Era has transformed modern biology, providing sequence data that records genetic changes that occur over time scales from billions of years (evolution) to months (tumor growth). This interdisciplinary course introduces the algorithms, statistics & biological concepts used to make inferences from genome datasets and will provide the computational foundation & practical experience needed to test biological questions using genome data.

Course was offered:  Spring 2018 · Spring 2017 · Spring 2016
BIOL 4240
History & Philosophy of Biol

This course will give an overview of the major conceptual and experimental advances in Biology. It will explore the relationships of Biology to mathematics and physical sciences and explore philosophical issues relevant to science in general, Biology in particular.

Course was offered:  Fall 2015 · Fall 2014 · Fall 2013
BIOL 4250
Human Genetics

Focuses on the fundamental knowledge about organization, expression, and inheritance of the human genome. Reviews classical Mendelian genetics and human genetic (pedigree) analysis. Emphasizes understanding human genetics in molecular terms. Includes gene mapping procedures, methodologies for identifying genes responsible for inherited diseases, the molecular basis of several mutant (diseased) states, the human genome project, and discussions about genetic screening and gene therapy. Prerequisite: BIOL 3010.

BIOL 4260
Cellular Mechanisms Offered Spring 2026

This course uses a case study approach to examine cellular processes that underlie diverse diseases and to identify the relevant molecular components that have been validated or that may serve as new therapeutic targets. We will discuss both established, transformative drugs as well as novel, emerging therapies under development. We will consider socio-economic and demographic issues that impact the accessibility and affordability of new drugs.

BIOL 4265
Developmental Neurobiology

The diverse functions of the nervous system depend on precise wiring of connections between neurons. This course covers cellular and molecular processes of how neuronal connections are established during development. Diseases which result from failing to establish the circuitry will also be discussed. This course will introduce research methods and technology, and encourage students to develop logical rationale of contemporary research.

Course was offered:  Fall 2019
BIOL 4270
Animal Behavior Laboratory

This laboratory course provides hands-on experiences with experimental approaches used to study animal behavior. The laboratory exercises explore visual and auditory sensory perception, biological clock, reproductive and aggressive behaviors using actively behaving animals such as hamsters, cichlid fish, crickets and electric fish. Students are given opportunities to design hypothesis-testing experiments in some laboratories.

BIOL 4280
Genetics of Behavior

This course studies behavior paradigms in model animals and the modern genetic tools used study and dissect the circuits underlying them. Can an animal as simple as a fly or mouse learn simple tasks, show appetitive behaviors and cravings, and inform studies of human addiction? Readings from classic and current literature will show the historical context of this field and develop critical reading skills. Prerequisites: BIOL 3000, BIOL 3010

BIOL 4310
Sensory Neurobiology Offered Spring 2026

This two-lectures-per-week course explores the basic principles of sensory neurobiology. The course consists of four modules. Each module represents one of the senses & consists of an introductory lecture, one or several lectures that will delve into the details of that sense, a current topic lecture on some recent finding, & finally, a guest lecture from a UVa researcher. Completion of BIOL 3050 or PSYC 2200 or PSYC 3200 strongly recommended.

BIOL 4320
Signal Transduction

This advanced undergraduate course explores how cells communicate with each other and respond to their environment. This area of biology is referred to as signal transduction and is the basis for most if not all normal and disease processes in humans. Therefore, significant time is spent on defining archetypal signaling modules that all cells use to receive and communicate information to and from their environment. Prerequisites: BIOL 3000 & BIOL 3010

BIOL 4330
Wiring the Brain Offered Spring 2026

This course focuses on how relatively simple model systems provide the clues as to how certain synaptic connections form and lead to specific behaviors. This will be followed by discussion of how this knowledge can be applied to the understanding and treatment of human neural disorders. 25% of the course is standard lectures and the rest, student-led discussion of primary literature. Prereqs: BIOL 3000 & BIOL 3010; BIOL 3050 or PSYC 2200 or 3200

BIOL 4340
Exper. Foundations of Neurobio

The course content will focus on three areas of neurobiological research: conduction of the nervous impulse, sensory physiology, and synaptic physiology. Prerequisites: Must have completed BIOL 3050 or BIOL 3170 or PSYC 4200

BIOL 4350
Metabolic Systems

A worldwide obesity epidemic exists. With it comes increased risk of chronic disease, such as Type 2 diabetes, cardiovascular disease, and certain cancers. This course will survey the molecular, genetic, physiologic and behavioral paths that lead to obesity and that contribute to prevalent chronic diseases. Through discussions of scientific literature, we will gain an integrated view of the factors that influence our energy homeostasis. Prerequiste: BIOL 3000, 3010.

Course was offered:  Spring 2018 · Spring 2017
BIOL 4360
Cytokine Signaling

This is a journal club format seminar where we perform an in depth analysis of the papers listed below. One paper will be covered per week with a review article also assigned for background. There are no presenters; rather we will have discussion leaders. All participants should be prepared to present any of the panels in the week's paper.

BIOL 4365
How to Map a Brain

If you want to understand how our brain works, this is the course for you! In this student-driven Journal Club-style seminar series, we will consider recent neuroscience literature for discussion of the most innovative discoveries. A broad range of outstanding neuroscience issues will be considered; topics could include, for example, strategies for gene therapy for human neurological diseases, or the remote control of learning and memory. Prerequisite: Instructor Permission

Course was offered:  Fall 2019 · Spring 2016 · Spring 2015
BIOL 4370
Epigenetics

Explores the emerging science, Epigenetics. Topics include epigenetics in model organisms and molecular mechanisms such as the Polycomb and Trithorax Group proteins, histone modifications and variants, dosage compensation, DNA methylation, nuclear reprogramming and stem cell pluripotency. Prerequisites: Genetics and Molecular Biology, Cell Biology and Biochemistry strongly recomended.

Course was offered:  Fall 2013
BIOL 4380
Evol Ecol Development

From the seahorse's body to the venus flytrap's jaws to the human brain, nature abounds with amazing adaptations. This interdisciplinary course explores how and why such biodiversity evolves as well as what limits diversity. Lectures and case studies will focus on core concepts, recent advances, and integrative approaches, placing special emphasis on the interplay between gene regulatory networks, the environment, and population genetics. Prerequisites: BIOL 3010, BIOL 3020

Course was offered:  Fall 2014
BIOL 4390
Biological Therapy of Cancer

This seminar course revolves around student-led presentations of primary literature in the field of cancer therapy using novel approaches including immunotherapies. Objectives include providing the student with significant exposure to primary literature and the development of critical thinking skills. Prerequisite: BIOL 3240.

BIOL 4430
Exptl Plant Biology Lab

We can't live without plants. Plants make our existence possible, and they hold secrets for a better future. Our experimental approach in this lab will combine genetics and genomics strategies to uncover some of those secrets. We'll search for genes and biosynthetic pathways that contribute to the success of plants at fighting off microbial infections. Ultimately, studies like these will lead to new, highly effective antimicrobial therapies. Prerequisite: BIOL 3010, BIOL 3150

BIOL 4440
Lipids and Membranes

Life requires lipids. Discussion of the literature will integrate lipids into our current protein-centric view of cell biology. Topics considered are current models of membrane structure and its effect on metabolism; synthesis and distribution of lipids to regulate cell communication, gene expression, and the coding of identity; how pathogens turn lipids against host cells; and how common pharmaceuticals affect lipid biology to treat disease.

Course was offered:  Spring 2017
BIOL 4450
Plant-Animal Interactions

Plants & animals have a long co-evolutionary history, with their interactions shaping natural ecosystems, as well as our own daily lives. We'll emphasize the evolutionary and ecological implications of these interactions to consider topics, such as pollination, herbivory and dispersal. We'll also address questions like: Why is flower color, shape and scent so diverse? How do animals eat toxic plants? How do fruit help plants finds new habitat?

Course was offered:  Fall 2020 · Fall 2019 · Fall 2018
BIOL 4460
Forest Sampling

Study of quantitative methods for sampling forest ecosystems

Course was offered:  Spring 2017 · Spring 2016 · Spring 2015
BIOL 4480
Macromolecular Structure

Exploration, in depth, of principles underlying protein and nucleic acid structures and the techniques used to determine those structures. Prerequisite: CHEM2410 and 2420 or BIOL3000 or permission of instructor

Course was offered:  Spring 2014
BIOL 4560
Crayfish neuro Lab

Course uses electrophysiological techniques with living crayfish material to examine principles of neurobiological function, including cellular resting potentials, propagated action potentials, neuromuscular physiology, aspects of neuromuscular organization, and sensory neuron physiology and organization. A lab lecture will precede each lab session. Grading will be based upon written laboratory reports and two midterm laboratory exams. Prerequisite: BIOL 3170

Course was offered:  Fall 2016 · Spring 2015 · Spring 2014
BIOL 4585
Selected Topics in Biology

Periodic seminar offerings to provide intensive study of the scientific literature in focused areas of Biology.

BIOL 4610
Molecular Evolution

Through the analysis of patterns of genetic variation in DNA, the field of Molecular Evolution seeks to gain insight into the fundamental evolutionary forces that generate, maintain, and remove genetic diversity. These forces shape the abundance of deleterious and beneficial mutations and reflect physical and behavioral differences between populations. In this course, we will dive into theoretical population genetics as a framework to develop an intuitive understanding of these evolutionary processes. We will apply this understanding to diversity among humans and all other life on earth. Prerequisite: BIOL3010, BIOL3020 required; STAT 2020 and calculus (MATH 1210, 1220, 1310, or 1320) suggested.

BIOL 4660
Biomedical Science Techniques Offered Spring 2026

How has a bioluminescent jellyfish saved lives? What does a Himalayan pond fish have to do with research into the origins of psychiatric disorders? Innovative methods in biomedical research have played a significant part in the development of revolutionary disease cures, treatments and diagnostics. This course will examine many of these technical approaches and how they have led to such significant discoveries in basic biomedical research. Prerequisite: BIOL 3010

BIOL 4751
Plant Diversity

The extraordinary diversity of the southern Appalachians will be used to explore the world of plants. We will visit unique mountain habitats to study the different species assemblages in these ecologically wide-ranging sites. Based upon our observations and analyses, we will critique contemporary views of the most effective conservation units (individual, population, species, family, habitat) and the methods used to achieve conservation goals.

BIOL 4752
Stream Ecology

Students will integrate principles of stream and watershed ecology to gain insight into stream-dwelling organisms and their environments. Participants will be introduced to the physical, chemical and biological organization of aquatic ecosystems, current theories in stream and watershed ecology, and lab and field methods for conducting stream research, and will participate in field/lab explorations and student-led discussions.

BIOL 4753
Biology of Fungi

The southern Appalachians provide an ideal setting to explore the biology of fungi. This class provides an introduction with emphasis on fieldID and current experimental methods used to study fungal genetics, ecology, and evolution. Lab exercises will use filamentous fungi to demonstrate methods for identification, culture techniques, breeding systems, genetic analysis, and interaction biology. Field trips will survey the taxonomic diversity.

Course was offered:  Summer 2016
BIOL 4754
Field Herpetology

We will focus on the ecology and evolution of reptiles and amphibians, leveraging their diversity in the southeastern US. In both the field and laboratory, we will study 1) the evolutionary relationships among reptiles and amphibians, 2) key evolutionary innovations that characterize each major lineage, 3) reptile and amphibian systems in ecological and evolutionary research, and 4) location and identification of reptiles and amphibians.

BIOL 4755
Field Biology of Fishes

MLBS sits on the Eastern Continental Divide providing an incredible diversity of freshwater habitats. Proficiency in ichthyology will be developed through field trips and lab work. Themes include: fish ID; patterns and drivers of diversity; interactions on individual, population, community and ecosystem levels; evolution; and influences of human activities. Students will design and conduct a research project and present at a class symposium.

BIOL 4757
GIS for Field Biologists

This course will cover the fundamentals of Geographic Information Systems as applied to biological questions with application in ecology, evolution, conservation, disease ecology, and human land-use. Students will learn spatial theory, analysis, and hands-on use of GIS software (including ArcGIS). Field laboratories will allow students to use Global Positioning Systems (GPS) and learn to incorporate this technology into spatial analyses.

Course was offered:  Summer 2014
BIOL 4758
Field Biology of Insects

Insects are perhaps the most important animal group on the face of the earth. Their enormous diversity makes them important models for understanding many concepts in biology. Students will observe the bits and pieces of an insect, they will discover how adaptation relates to diversity, and they will learn to identify the major insect groups. Field trips to varied habitats allow students to collect insects and understand their natural history.

Course was offered:  Summer 2025 · Summer 2019 · Summer 2017
BIOL 4759
Field Mtds in Wildlife Ecol

An introduction to field research methods for measuring and monitoring animals with an emphasis on testing biological and wildlife management hypotheses. We will survey small mammals, birds, reptiles and amphibians. Students will learn sampling designs, protocols, and types of studies. Exercises will include surveying, trapping, marking, and measuring animals. Skills learned will be used in hypothesis-driven group projects.

Course was offered:  Summer 2015
BIOL 4760
Hormones and Behavior

Hormones alter the development and expression of animal behavior. Behavior in turn changes the effects of hormones. We'll take an evolutionary approach in exploring the causation and mechanism of hormone-mediated behaviors. We will use endocrinological techniques to examine behavior and hormone variation in wild populations. Students will help design and conduct a class research project with the goal of publishing our results.

Course was offered:  Summer 2015
BIOL 4761
Wildlife Disease Ecology

This course focuses on the ecology and evolution of infectious diseases in wild animal populations. Topics include the population biology of parasites and pathogens, host immune defenses and pathogen virulence, and wildlife conservation and disease. Students will gain experience with quantitative methods and field and laboratory techniques, including parasite identification and handling of insects, birds, amphibians, and small mammals.

Course was offered:  Summer 2024 · Summer 2018
BIOL 4762
Field Behavioral Ecology

This course will illustrate principles of behavior and provide experience with methods used in animal behavior research. Students will develop an understanding of the scientific process as applied to behavior research, learn how behavior evolves and why we see the behavioral patterns that we do, and learn how to conduct research in wild populations. The class will work collaboratively to develop and carry out a field research project.

Course was offered:  Summer 2023 · Summer 2022
BIOL 4770
Synthetic Biology Offered Spring 2026

By applying the principles of engineering to biology, students will design molecules, viruses, and cells to solve global problems in public health, food security, manufacturing, information processing, and the environment, changing the traditional question of 'How do cells work?' to 'How can I get a cell to work for me?' Students will gain experience in writing internationally competitive research project proposals. Prerequisite: Instructor Permission

BIOL 4810
DMP Seminar- Biology Research

Two-hour, weekly discussion of recent advances in biology; attend biology seminars, interact with seminar speakers, explore the philosophy and practice of science, and learn skills in oral and written research presentation. Prerequisite: Fourth-year DMP in Biology.

BIOL 4820
DMP Seminar- Biology Research Offered Spring 2026

Two-hour, weekly discussion of recent advances in biology; attend biology seminars, interact with seminar speakers, explore the philosophy and practice of science, and learn skills in oral and written research presentation. Prerequisite: Fourth-year DMP in Biology.

BIOL 4840
BS Capstone Biology

This course will explore paths towards careers in biology and the skills and training required to pursue them. The course will include visits from professional biologists and UVA alumnae, who will provide advice on navigation of their careers in biology. The course will also offer guidance on additional training, advanced degrees, and internships. Skills are cultured by scientific abstracts, peer and mentor interaction, and interest statements. Prerequisites: BIOL 3000, 3010 and 3020. Students who have not completed these courses may be able to take them concurrently with permission.

Course was offered:  Fall 2025
BIOL 4900
Independent Study in Biology Offered Spring 2026

Independent study under the direction of a Biology faculty member for students to read and critically assess primary research papers and current reviews in a focused area of the life sciences. Directed readings and discussions can be used to explore how contemporary topics and research areas can be incorporated into other formal courses. Students will have the opportunity to develop both scientific writing and oral presentation skills.

BIOL 4910
Life Sciences Research Offered Spring 2026

Undergraduate research under the direction of a UVA Professor who doesn't belong to the Biology Department. Despite the 'Closed' status of BIOL4910 on SIS, the course is open for enrollment. For application instructions, see the section 'How to Enroll in Independent Research with a Faculty Member Outside of the Biology Department' at: https://bio.as.virginia.edu/undergraduate/research. Prerequisite: Instructor Permission

BIOL 4911
Independent Research I

Independent research for qualified undergraduates under the direction of a faculty member. Prerequisite: Instructor permission.

Course was offered:  Summer 2014 · Spring 2014 · Fall 2013
BIOL 4912
Independent Research II

Independent research for qualified undergraduates under the direction of a faculty member. Prerequisite: Instructor permission and BIOL 4911.

Course was offered:  Summer 2014 · Spring 2014 · Fall 2013
BIOL 4913
Independent Research III

Independent research for qualified undergraduates under the direction of a faculty member. Prerequisite: Instructor permission and BIOL 4912.

Course was offered:  Summer 2014 · Spring 2014 · Fall 2013
BIOL 4914
Independent Research IV

Independent research for qualified undergraduates under the direction of a faculty member. Prerequisites: Instructor Permission and BIOL 4913.

Course was offered:  Summer 2014 · Spring 2014 · Fall 2013
BIOL 4915
Independent Research V

Independent research for qualified undergraduates under the direction of a faculty member. Prerequisite: Instructor permission and BIOL 4914.

Course was offered:  Summer 2014 · Spring 2014 · Fall 2013
BIOL 4916
Independent Research VI

Independent research for qualified undergraduates under the direction of a faculty member. Prerequisite: instructor permission and BIOL 4915.

Course was offered:  Summer 2014 · Spring 2014 · Fall 2013
BIOL 4917
Independent Research VII

Independent research under the guidance of a departmental faculty member. Prerequisite: Instructor permission and BIOL 4916.

Course was offered:  Summer 2014 · Spring 2014 · Fall 2013
BIOL 4918
Independent Research VIII

Independent research under the guidance of a departmental faculty member. Prerequisite: Instructor Permission and BIOL 4917

Course was offered:  Summer 2014 · Spring 2014 · Fall 2013
BIOL 4930
Distinguished Major Research Offered Spring 2026

This course is the final semester of Independent Research for participants of the Biology Distinguished Majors Program. During this semester, students will complete their laboratory investigations, ultimately presenting the sum of their work in a written thesis. Prerequisite: Instructor Permission

BIOL 4940
BS Capstone Research Offered Spring 2026

This course will provide students with essential hands-on experience in experimental design, data collection, analysis, and science communication. Students will have the opportunity to think critically and creatively about biology and to develop research competencies. Students will work with a mentor to develop a project that makes a unique scientific contribution and will communicate their research to a broader audience at the University's Undergraduate Research Symposium or similar.

Course was offered:  Spring 2026
BIOL 4993
Independent Study In Biology

Independent study under the direction of a Biology faculty member for students to read and critically assess primary research papers and current reviews in a focused area of the life sciences. Directed readings and discussions can be used to explore how contemporary topics and research areas can be incorporated into other formal courses. Students will have the opportunity to develop both scientific writing and oral presentation skills.

BIOL 4994
Ind Study in Biology

Independent study for qualified undergraduates under the direction of a faculty member. Prerequisite: Instructor permission and BIOL 4993.

BIOL 4995
Independent Study

Independent study for qualified undergraduates under the direction of a faculty member.

Course was offered:  Spring 2015 · Spring 2014
BIOL 5010
Biochemistry

Biochemistry underlies nearly every biological process, from environmental science to medicine. When living systems are in chemical and energetic balance, organisms thrive. When they're out of balance, as in disease or unpredictable environments, life is compromised. This course will explain how simple chemical and physical principles apply to the major classes of biological macromolecules that maintain life. Prerequisite: BIOL 2010 or BIOL 2100 or BME 2104 and BIOL 2020 or BIOL 2040 and either CHEM 2410 or CHEM 1820

Course was offered:  Summer 2014 · Fall 2013
BIOL 5070
Prac Aspects Light Microscopy Offered Spring 2026

Practical usage of various microscopy imaging methodologies to study the morphology and cellular function in various biological systems from single cell to single molecule in cells and tissues. Topics include basics theory of microscopy, imaging and image analysis to solve various biological questions, fluorophore labeling, technical and hands on training on various microscopy techniques applied in different biological and biomedical investigations. Lectures, discussion, student presentations and laboratory.

BIOL 5559
New Course: BIOL

New course in the subject of biology.

Course was offered:  Summer 2016
BIOL 7013
Stem Cells

The course will deep dive into what stem cells are, what they do, where and how they function, and how we can use stem cells in the clinic to repair damaged tissue and restore tissue function. The course will consist of a series of lectures and student run discussions related to current scientific literature.

Course was offered:  Spring 2022
BIOL 7030
Computational Evol Biology Lab

The evolutionary history of a population can be studied by examining patterns of genetic variation among individuals from a species. In this lab course, you will learn how to utilize genomic data to make evolutionary inferences. Together, we will learn fundamentals of population genetics, how to conduct bioinformatic research on high-performance computers, how to take raw sequence data and turn it into biological insight, and how to present the results of your research.

Course was offered:  Fall 2025
BIOL 7045
Neurodegenerative Diseases

This course will focus mainly on research about Alzheimer's disease, and will meet once a week for 3 hours The first 3 weeks will be primarily didactic, and the remainder of the course will be a "journal club" in which primary research paper discussions will be led by teams of students. Assessments will be based on how well students lead and participate in discussions, and on exams.

Course was offered:  Fall 2022
BIOL 7060
Organ Dev Tissue Eng

Why do most of our adult body tissues have limited regenerative capacity? How can terminally diseased organs be replaced? This course will cover the cellular mechanisms that regulate animal tissue formation, regeneration and repair in vivo. Students will gain insights into the opportunities, limitations, and risks of tissue engineering in vitro, as an emerging research field that may lead to revolutionary organ replacement strategies. Prerequisite: BIOL 3000

Course was offered:  Spring 2015
BIOL 7110
Teaching Science

This STEM teaching course will help graduate TAs integrate learning theory and effective student engagement practices into their teaching. GTAs will participate in guided discussions to relate recommendations from the education literature to their classroom experiences. Assignments will include learning activities, such as teaching observations & reflections, and designing interventions to assist students with difficult topics/skills.

BIOL 7120
When Good Cells Go Bad

This course will cover topics related to major neurodegenerative diseases including Multiple Sclerosis (MS), Muscular Dystrophy (MD), Peripheral Nerve Sheath Tumor (Neurofibrosarcoma) and Malignant peripheral nerve sheath tumor (MPNST). Topics related to pathology and molecular mechanism of diseases, possible drug discovery targets, and therapeutic discovery approaches will be emphasized.

Course was offered:  Fall 2014 · Fall 2013
BIOL 7140
NextGen Sequencing and Apps

Students will learn the next generation sequencing technologies and explore their applications in the studies of evolution and ecology. This course is a lecture and journal club format where primary scientific literature will be discussed. Students will also learn basic bioinformatic skills.

Course was offered:  Fall 2013
BIOL 7150
Evolution of Sex

Despite the many benefits of asexual reproduction, the vast majority of eukaryotic organisms reproduce sexually. How sex evolved, and how it persists despite its many associated costs, are major unanswered questions in biology. We will explore the diversity of sexual reproduction and associated evolutionary phenomena with a focus on critically evaluating current research and theory in this field. Prerequisite: BIOL 3020 or permission from Instructor

BIOL 7180
Behavioral Ecology

Behavioral ecology explores the evolutionary analysis and explanations for the diversity of animal behavior, including foraging decisions, altruism, cooperation, mate choice, group living, parental care and range of other sociobiological phenomena.

Course was offered:  Fall 2014 · Fall 2013
BIOL 7190
Biological Clocks

Introduces biological timekeeping as used by organisms for controlling diverse processes, including sleep-wakefulness cycles, photoperiodic induction and regression, locomotor rhythmicity, eclosion rhythmicity, and the use of the biological clock in orientation and navigation.

Course was offered:  Fall 2015 · Fall 2014 · Fall 2013
BIOL 7220
Systems Biology

An introduction to a new research paradigm that focuses on the systematic study of complex interactions at the molecular, network and genomic level. This course will review state-of-the-art high throughput techniques and modeling methods used to obtain, integrate and analyze complex data from biological systems. This course will be a combination of text based lectures and discussions of the current literature pertinent to Systems Biology.

Course was offered:  Spring 2014
BIOL 7230
Bioinformatics

The Genome Era has transformed modern biology, providing sequence data that records genetic changes that occur over time scales from billions of years (evolution) to months (tumor growth). This interdisciplinary course introduces the algorithms, statistics & biological concepts used to make inferences from genome datasets and will provide the computational foundation & practical experience needed to test biological questions using genome data.

Course was offered:  Spring 2018 · Spring 2017 · Spring 2016
BIOL 7240
Molecules to Mind

This intensive, graduate-level course is designed to provide a foundational understanding of the principles underlying the development, genetics, and molecular-cellular biology of the nervous system. Over the span of the semester, students will engage with three core areas of neuroscience: Genetics in Neuroscience, Neurodevelopment, Molecular and Cellular Biology (MCB) of Neurons.

Course was offered:  Fall 2025 · Fall 2024 · Fall 2022
BIOL 7280
Genetics of Behavior

This course studies behavior paradigms in model animals and the modern genetic tools used study and dissect the circuits underlying them. Can an animal as simple as a fly or mouse learn simple tasks, show appetitive behaviors and cravings, and inform studies of human addiction? Readings from classic and current literature will show the historical context of this field and develop critical reading skills.

BIOL 7310
Sensory Neurobiology

This two-lectures-per-week course explores the basic principles of sensory neurobiology. The course consists of four modules. Each module represents one of the senses and consists of an introductory lecture, one or several lectures that will delve into the details of that sense, a current topic lecture on some recent finding, and finally, a guest lecture from a UVa researcher. Prerequisites: Instructor Permission.

BIOL 7320
Signal Transduction

This advanced undergraduate course explores how cells communicate with each other and respond to their environment. This area of biology is referred to as signal transduction and is the basis for most if not all normal and disease processes in humans. Therefore, significant time is spent on defining archetypal signaling modules that all cells use to receive and communicate information to and from their environment.

BIOL 7360
Cytokine Signaling

This is a journal club format colloquium where we perform an in depth analysis of the papers listed below. One paper will be covered per week with a review article also assigned for background. There are no presenters; rather we will have discussion leaders. All participants should be prepared to present any of the panels in the week's paper.

BIOL 7370
Epigenetics

Explores the emerging science, Epigenetics. Topics include epigenetics in model organisms and molecular mechanisms such as the Polycomb and Trithorax Group proteins, histone modifications and variants, dosage compensation, DNA methylation, nuclear reprogramming and stem cell pluripotency.

Course was offered:  Fall 2013
BIOL 7380
Evo Eco Devo

From the seahorse's body to the venus flytrap's jaws to the human brain, nature abounds with amazing adaptations. This interdisciplinary course explores how and why such biodiversity evolves as well as what limits diversity. Lectures and case studies will focus on core concepts, recent advances, and integrative approaches, placing special emphasis on the interplay between gene regulatory networks, the environment, and population genetics. Prerequisite: BIOL 3010, BIOL 3020

Course was offered:  Fall 2014
BIOL 7559
New Course: BIOL

New course in the subject of biology.

BIOL 7660
Biomedical Science Techniques

Do you know how a bioluminescent jellyfish protein is saving lives? The green fluorescent protein, earning its discoverers the 2008 Nobel Prize, is only one example of the recent biomedical breakthroughs leading to revolutionary diagnostics, treatments and cures that we will cover. Topics will range from how scientists are using roundworms to cure diabetes to why a pond fish from Himalayas might unlock the mysteries of psychiatric disorders.

Course was offered:  Fall 2018 · Fall 2017
BIOL 7751
Plant Diversity

The extraordinary diversity of the southern Appalachians will be used to explore the world of plants. We will visit unique mountain habitats to study the different species assemblages in these ecologically wide-ranging sites. Based upon our observations and analyses, we will critique contemporary views of the most effective conservation units (individual, population, species, family, habitat) and the methods used to achieve conservation goals.

BIOL 7752
Stream Ecology

We will focus on integrating principles of stream and watershed ecology to gain insight into stream dwelling organisms and their environments. Students will be introduced to 1) the physical, chemical and biological organization of aquatic ecosystems, 2) current theories in stream and watershed ecology, and 3) lab and field methods for conducting stream research. Students will conduct independent and group research projects.

Course was offered:  Summer 2014
BIOL 7753
Field Biology of Fungi

The southern Appalachians provide an ideal setting to explore the biology of fungi. This class provides an introduction with emphasis on fieldID and current experimental methods used to study fungal genetics, ecology, and evolution. Lab exercises will use filamentous fungi to demonstrate methods for identification, culture techniques, breeding systems, genetic analysis, and interaction biology. Field trips will survey the taxonomic diversity.

Course was offered:  Summer 2016
BIOL 7754
Field Herpetology

We will focus on the ecology and evolution of reptiles and amphibians, leveraging their diversity in the southeastern US. In both the field and laboratory, we will study 1) the evolutionary relationships among reptiles and amphibians, 2) key evolutionary innovations that characterize each major lineage, 3) reptile and amphibian systems in ecological and evolutionary research, and 4) location and identification of reptiles and amphibians.

BIOL 7755
Field Biology of Fishes

MLBS sits on the Eastern Continental Divide providing an incredible diversity of freshwater habitats. Proficiency in ichthyology will be developed through field trips and lab work. Themes include: fish ID; patterns and drivers of diversity; interactions on individual, population, community and ecosystem levels; evolution; and influences of human activities. Students will design and conduct a research project and present at a class symposium.

BIOL 7757
GIS for Field Biologists

This course will cover the fundamentals of Geographic Information Systems as applied to biological questions with application in ecology, evolution, conservation, disease ecology, and human land-use. Students will learn spatial theory, analysis, and hands-on use of GIS software (including ArcGIS). Field laboratories will allow students to use Global Positioning Systems (GPS) and learn to incorporate this technology into spatial analyses.

Course was offered:  Summer 2014
BIOL 7758
Field Entomology

Insects are perhaps the most important animal group on the face of the earth. Their enormous diversity makes them important models for understanding many concepts in biology. Students will observe the bits and pieces of an insect, they will discover how adaptation relates to diversity, and they will learn to identify the major insect groups. Field trips to varied habitats allow students to collect insects and understand their natural history.

BIOL 7759
Wildlife Techniques

An introduction to field research methods for measuring and monitoring animals with an emphasis on testing biological and wildlife management hypotheses. We will survey small mammals, birds, reptiles and amphibians. Students will learn sampling designs, protocols, and types of studies. Exercises will include surveying, trapping, marking, and measuring animals. Skills learned will be used in hypothesis-driven group projects.

Course was offered:  Summer 2015
BIOL 7760
Hormones and Behavior

Hormones alter the development and expression of animal behavior. Behavior in turn changes the effects of hormones. We'll take an evolutionary approach in exploring the causation and mechanism of hormone-mediated behaviors. We will use endocrinological techniques to examine behavior and hormone variation in wild populations. Students will help design and conduct a class research project with the goal of publishing our results.

Course was offered:  Summer 2015
BIOL 7761
Wildlife Disease Ecology

This course focuses on the ecology and evolution of infectious diseases in wild animal populations. Topics include the population biology of parasites and pathogens, host immune defenses and pathogen virulence, and wildlife conservation and disease. Students will gain experience with quantitative methods and field and laboratory techniques, including parasite identification and handling of insects, birds, amphibians, and small mammals.

Course was offered:  Summer 2024 · Summer 2018
BIOL 7762
Field Behavioral Ecology

This course will illustrate principles of behavior and provide experience with methods used in animal behavior research. Students will develop an understanding of the scientific process as applied to behavior research, learn how behavior evolves and why we see the behavioral patterns that we do, and learn how to conduct research in wild populations. The class will work collaboratively to develop and carry out a field research project.

Course was offered:  Summer 2023 · Summer 2022
BIOL 7993
Independent Study in Biology

A biology faculty member supervises and approves all components of this course, designating the number of credits to be earned prior to enrollment. Students successfully complete one or more courses offered by the Department of Biology at the 3000 level or above and, for each course, write a 10-page (minimum) paper on a relevant topic.

BIOL 7994
Independent Study in Biology Offered Spring 2026

Independent research for qualified graduates under the direction of a faculty member within the Biology Department.

BIOL 8010
Integrative Biology Colloquium

A weekly conference in which students present reports covering various aspects of Integrative Biology. May be repeated for credit.

BIOL 8011
Integrative Biology I

Integrative Biology seeks to understand how genetic, cellular, and developmental processes give rise to organismal phenotypes and how changes in these processes over time contribute to the evolution of form and function. In this course, graduate students will learn principles of genetics, cell, developmental, and organismal biology and apply this knowledge towards answering fundamental questions in biology.

Course was offered:  Fall 2025
BIOL 8050
Adv. Evolutionary Biol

This course will cover a range of evolutionary concepts and approaches, including levels of selection, the role of evolution in structuring ecological communities, game theoretical models of adaptation, frequency-dependence, neutral processes and drift, the evolution of sex, the evolution of virulence, the molecular basis of adaptation, population and quantitative genetics, and the evolution of genome structure.

Course was offered:  Fall 2024
BIOL 8060
Colloquium: Circadian Biology

Readings and two-hour student seminar preparations focusing on recent research and primary literature in circadian biology. Prerequisites: Instructor Permission

Course was offered:  Fall 2015 · Fall 2014 · Fall 2013
BIOL 8070
Colloquium in Population Biol Offered Spring 2026
BIOL 8080
Colloq in Fundam Neuroscience Offered Spring 2026

This weekly colloquium provides a forum for exploring cutting-edge research in fundamental neuroscience. The course format includes journal club discussions of recent high-impact publications, as well as presentations of ongoing research by both students and faculty members.

Course was offered:  Spring 2026 · Spring 2025
BIOL 8081
Advanced Ecology & Evolution 1

This course introduces grad students to a breadth and depth of concepts and theories in modern ecology and evolutionary biology. The couse is co-taught by two BIOL faculty each fall, with different faculty rotating into the course in alternate years, providing expertise in molecular population genetics, genomics, phylogenetics, integrative biology, speciation, microevolution, life-history evolution, and mating systems.

BIOL 8082
Advanced Ecology & Evolution 2 Offered Spring 2026

This course introduces grad students to a breadth and depth of concepts and theories in modern ecology and evolutionary biology.. The course is taught by a different BIOL faculty each spring, with different faculty rotating into the course in alternate years, providing expertise in molecular population genetics, genomics, phylogenetics, integrative biology, speciation, microevolution, life-history evolution, and mating systems.

BIOL 8083
Advanced Ecology & Evolution 3

This course introduces grad students to a breadth and depth of concepts and theories in modern ecology and evolutionary biology. The couse is co-taught by two BIOL faculty each fall, with different faculty rotating into the course in alternate years, providing expertise in molecular population genetics, genomics, phylogenetics, integrative biology, speciation, microevolution, life-history evolution, and mating systems.

BIOL 8084
Advanced Ecology & Evolution 4 Offered Spring 2026

This course introduces grad students to a breadth and depth of concepts and theories in modern ecology and evolutionary biology.. The course is taught by a different BIOL faculty each spring, with different faculty rotating into the course in alternate years, providing expertise in molecular population genetics, genomics, phylogenetics, integrative biology, speciation, microevolution, life-history evolution, and mating systems.

BIOL 8240
Professional Skills I

Introduces professional skills to first-year graduate students in the life sciences. This includes personal skills such as research management, ethics training and career awareness. Skills for communicating in science include small grant writing, poster creation and presentation. Emphasis will be placed on developing a scientific community that recognizes and values diversity.

BIOL 8250
Prof Skills Life Sciences II Offered Spring 2026

Introduces professional skills to first-year graduate students in the life sciences. This includes personal skills such as time management, ethics training and professional identity. Skills for communicating in science include oral presentations to a variety of audiences. Emphasis will be placed on developing a scientific community that recognizes and values diversity.

BIOL 8260
Writing in Science

Developing skill in communicating scientific principles and writing compelling research proposals is essential for successful graduate training in the biological sciences. This seminar and workshop course will focus on how to create effective grant and research proposals in preparation for thesis research. Students will be actively involved by presenting their research progress and plans, and critiquing each other's written proposals.

BIOL 8270
Sev Hab Hig Eff Grd Stds

Introduces professional skills to first-year graduate students in the life sciences. This includes personal skills such as research management, ethics training and career awareness. Skills for communicating in science include small grant writing, poster creation and presentation. Emphasis will be placed on developing a scientific community that recognizes and values diversity.

BIOL 8300
Studies of the Phenotype

This course will explore case studies of research on phenotypes that emphasize the insights and advances gained by bringing a multilevel integrative approach to a target question. Students will learn to apply an integrative perspective to their own research topics by developing a research proposal that expands across levels. This class is open to all life science PhD students with recommendation for those in their 1st or 2nd year.

Course was offered:  Fall 2025 · Fall 2024 · Fall 2023
BIOL 8310
Career Design

This course uses a student-centered approach to explore the diverse careers available to life science PhD students. Students will identify their skills and experiences that match with professional opportunities, and they will develop a career individual development plan to guide their career planning. This class is open to all life science PhD students after the 2nd year in their program. This class is part of the PhD Plus Career Design series.

Course was offered:  Fall 2025 · Fall 2024 · Fall 2023
BIOL 8320
Professional Transitions

Students will develop the necessary skills and realistic plans for finishing their dissertations and acquiring a position. Website, curriculum vitae, cover letter, research description, teaching portfolio, and summary of professional experience will be developed. This class is open to all life science PhD students in their 4th or 5th year.

Course was offered:  Fall 2023
BIOL 8410
Little Cells, Big Data Offered Spring 2026

This course will integrate lectures and workshops to teach students how to analyze publicly available single-cell 'omic datasets relevant to their research interests, which students will perform as part of a scaffolded project. The course will cover a variety of methods, with a focus on single-cell RNA-seq.

Course was offered:  Spring 2026
BIOL 8559
New Course: BIOL

New course in the subject of biology.

BIOL 8840
Selected Topics in Physiology

A discussion of current problems.

Course was offered:  Spring 2021 · Spring 2020 · Spring 2019
BIOL 8870
Sel Top in Develpmntl Genetic

A discussion of current problems. Prerequisite: Instructor permission.

BIOL 8998
Non-Topical Rsch, Masters Prep

For master's research, taken before a thesis director has been selected.

BIOL 9559
New Course: BIOL

New course in the subject of biology.

Course was offered:  Fall 2015
BIOL 9910
Rotation Research

An exposure to the working techniques and interactions of the modern Biological Laboratory. Required of all first-year biology graduate students.

BIOL 9920
Rotation Research Offered Spring 2026

An exposure to the working techniques and interactions of the modern Biological Laboratory. Required of all first-year biology graduate students.

HBIO 4810
Capstone Sem Human Biology I

A weekly seminar co-organized by participating faculty to integrate students' independent research and coursework with contemporary issues at the intersection of biology, the humanities and social sciences. Students will have the opportunity to present their ongoing research and meet with outside speakers. This course will be taken in the fourth year. Prerequisite: DMP in Human Biology.

HBIO 4820
Capstone Sem Human Biology II Offered Spring 2026

A weekly discussion and workshop co-organized by participating faculty to provide guidance and advice to students on completing their research or independent study and writing their thesis. Occasional seminars and opportunities to meet outside speakers will continue in this semester. This course will be taken in the fourth year. Prerequisite: DMP in Human Biology.

HBIO 4950
Indep Research for Human Biol

Independent research/independent study under the guidance of a primary mentor within the College of Arts and Sciences. Prerequisite: DMP in Human Biology.

HBIO 4960
Indep Research for Human Biol Offered Spring 2026

Independent research/independent study under the guidance of a primary mentor within the College of Arts and Sciences. Prerequisite: DMP in Human Biology.

HBIO 4998
Thesis Rsrch in Human Biol I

Independent research/independent study under the guidance of a primary mentor within the College of Arts and Sciences. Research/study forms the basis for the DMP thesis to be submitted at the end of the fourth year. This course must be taken in the first semester of the fourth year and should encompass the majority of the research for the thesis. Prerequisite: First-semester fourth-year DMP in Human Biology.

HBIO 4999
Thesis Rsrch in Human Biol II Offered Spring 2026

This course is designed to provide students the opportunity for hands-on learning in experimental sciences leading to a Capstone thesis project and written thesis. Students, working with a primary mentor (and in some cases a secondary mentor), design an original research study or other creative product in self-selected areas of interest, execute the study, analyze the data and report the findings in written form.